Department of Biology
ESC 350
200 University Ave. W
Waterloo, Ontario, Canada N2L 3G1
Phone: (519) 888-4567 ext. 32569
Fax: (519) 746-0614
Prairie and Montane Goldenasters and Telegraph Weeds
Heterotheca Cassini includes all the goldenasters that have leaf hairs consisting of bone-shaped (osteolate) cells with knobbed surfaces.
The genus is divided into three sections based on fruit characteristics, presence or absence of ray florets, and growth form. Species of Heterotheca are native to southwestern Canada, much of the U.S. and most of Mexico.
Heterotheca sect. Heterotheca
Heterotheca sect. Heterotheca includes three species with heterocarpic heads (two kinds of fruit; e.g., H. subaxillaris). The two species native to the U.S. are weedy annuals sometimes reaching 2 m in height, but can be short lived perennials in Mexico
- Heterotheca grandiflora is an annual native to California
- Heterotheca inuloides is a perennial native to Mexico.
- Heterotheca subaxillaris is native to the central and southern Great Plains eastward to New York and south to Florida and into the mountains Mts into northern Mexico.
Heterotheca sect. Ammodia
Heterotheca sect. Ammodia includes one species H. oregona with four varieties native to gravel bars and arroyos from Washington to Central California. The species has rayless heads.
Heterotheca sect. Phyllotheca
Heterotheca sect. Phyllotheca (Nutt.) Harms (Semple 1996) includes 19 species of mostly herbaceous perennials ranging in height from a few cm (H. jonesii) to more than 1.2 m (H. shevockii). Most species are usually 10-50 cm tall (e.g., H. pumila). Many species are restricted to particular soil types or habitats (e.g., H. brandegei, Baha CA Norte; H. monarchensis, CA; a few species are widely distributed (e.g., H. villosa; nine varieties). Species are defined by different combinations of leaf traits (size, shape, numbers of hairs and glands), head features (size, numbers of florets) and habit (growth form) features. All line drawing illustrations are from Semple (1996) and are © J.C. Semple (1996).
- Heterotheca barbata
- Heterotheca brandegei
- Heterotheca camporum
- Heterotheca canescens
- Heterotheca fulcrata
- Heterotheca gypsophila
- Heterotheca jonesii
- Heterotheca monarchensis
- Heterotheca marginata
- Heterotheca mexicana
- Heterotheca mucronata
- Heterotheca pumila
- Heterotheca rutteri
- Heterotheca sessiliflora
- ssp. bolanderi
- ssp. echioides
- ssp. fastigiata
- ssp. sessiliflora
- Heterotheca shevockii
- Heterotheca stenophylla
- Heterotheca thiniicola
- Heterotheca villosa
- var. ballardii
- var. depressa
- var. foliosa
- var. nana (including var. horrida)
- var. minor (including var. hispida)
- var. pedunculata
- var. scabra
- var. sierrablancensis
- var. villosa
- Heterotheca viscida
- Heterotheca zionensis
Last update 31 October 2013 by J.C. Semple
© 2013 J.C. Semple, including all photographs unless otherwise indicated